#lions#wildlifedocumentary#wildlife In Botswana’s Okavango Delta, two brothers win a ferocious battle to rule a large lion pride. Their arrival brings security, and five of the six lionesses soon give birth to thirteen cubs. Lions in Battle follows the fast-growing Xakanaxa pride through one of the Delta’s harshest winters. With twenty-one mouths to feed, the lions target buffalo, but when the herds move beyond Xakanaxa into enemy territory, their main food source vanishes almost overnight. The sixth lioness’s late cubs struggle to keep pace, and starvation sets in.
In desperation, the lionesses turn on an elephant herd, killing six within a month as the elephants fight back with tactics of their own. When the summer rains bring the buffalo home, the pride feasts again.
They have won this battle, but in the unpredictable Okavango, their future is never certain.
0:00 The Unforgiving Okavango Delta 4:49 Rules of Engagement 8:15 The Turning Season 12:39 Lions vs Buffalo 18:06 More Lions, Less Space 23:41 Desperation for the Lion Pride 28:05 Lions Hunt Elephants 34:00 The Return of the Rains 40:34 Lion Pride Test Strength Against Buffalo
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Video Summary
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A lion pride in Botswana's Okavango Delta faces a brutal cycle of prosperity and starvation. After a baby boom, the migration of their primary prey, cape buffalo, leads to devastating cub losses. The pride survives by adapting their hunting tactics to target elephants, eventually regaining stability as the rains return and the buffalo herds migrate back to the region.
Ever-changing, ever-challenging, the Okavango Delta
in Southern Africa is as unforgiving as the creatures that live here.
One successful lion pride, led by two brothers, sees
its fortune reverse as the Okavango deals out one
of its harshest seasons ever.
In their struggle to survive,
these lethal predators face off against well-armed prey.
Things only get worse.
Pushed to the brink of starvation,
the pride will resort to hunting even more difficult prey,
forced by fickle circumstances
to adapt or die.
In the heart
of Botswana's
Okavango Delta
spreads a lush area of islands and floodplains.
This area, called Kakonika,
is a summer paradise for all its inhabitants,
including its lions.
Resident pride has been unsettled,
Torn by a battle for power between different males
for the right to dominate.
And so, the six females have not given birth for over a year.
By the beginning of summer,
two powerful brothers win the right to rule the Kakonika pride,
bring stability to the pride,
and over the next six months,
the lionesses give birth one after the other.
In December, two females have five cubs.
Three months later, another three lionesses deliver eight cubs.
In such a short period of time,
the pride increases by a total of 13 cubs.
In the secluded dens,
the little ones play during their few waking hours.
Each female keeps her own nursing cubs isolated
from the pride for the first two months.
Born at the height of summer, these cubs,
the future of the pride, are off to a good start.
But this unusual bounty of healthy cubs,
once they've been weaned,
could cause problems in the long term.
At three months old, they start to eat meat.
And all of a sudden,
the adults have an extra 13 mouths to feed.
To ensure the cubs' well-being,
the mothers have to provide meat at regular intervals,
every three
or four days.
Welcomed into the pride,
the cubs thread their way
into the social fabric.
Their life skills are sharpened by their fathers.
And the games they play with each other.
The Kakonika brothers are extremely patient and tolerant fathers.
The lionesses form the core of the pride,
bound by one collective purpose, to hunt.
In summer, they hunt large cape buffalo,
abundant in this part of the Okavango,
but only for half the year.
When the rains stop and the buffalo deplete the grasses, they'll leave.
But while they feast, so do the lions.
Each herd has
400 or more
buffalo.
perfect balance of numbers where
the births outweigh the killings by the
lions. They are one of the most desirable
and dangerous of prey. Desirable because
the meat from a one-ton Buffalo can feed
the entire pride for a week. Dangerous
because their formidable horns can inflict
a fatal wound and if the herd
stampedes the Lions can't outrun them.
But the six Kakonika lionesses learned
They learned all this early in their hunting careers.
They know the rules of engagement.
They single out a buffalo,
one they can separate from the herd.
This time,
they choose a mother,
concentrating their power
on bringing her down.
They ignore her newborn calf.
For the six-month-old cubs,
the calf presents a learning opportunity.
Every chance they get to practice hunting
brings them closer to independence.
By the age of two, they will hunt on their own.
They're fast learners,
every death is a way of life.
And each kill is a step forward in their own survival.
The youngest cubs get led to the kill by their mother.
At three months old,
they find the mountain of meat both tasty and entertaining.
At each meal, adults and cubs gorge themselves,
feeding side by side.
This is unusual. Lions normally eat according to hierarchy.
Males first, cubs last.
Perhaps the abundance of food allows
the Kakanaka pride to relax the rules.
Though the youngsters still get scolded to mind their manners.
During summer,
while the buffalo roam
the Kakanaka territory, the pride
is guaranteed regular meals.
By May, the end of summer,
the rains stop
and the lion's good
fortune starts to change.
Breeding herds of elephants herald this shift in season.
They arrive from the surrounding Kalahari Desert and spread
into the forests and flood plains in the heart of the Okavango.
They've come because the water in the surrounding desert has dried,
and the vegetation has shriveled.
Kakanaka offers them permanent water.
The elephants need between 20
and 40 gallons each every day.
Remarkably, they can drink that in just five minutes
by filling their trunks and squirting
it down their throats.
Lack of water is the biggest
killer of elephants,
more so than starvation.
The beginning of winter marks the start
of hardship for the large Kakanaka pride.
For now, the cubs are oblivious of the challenges to come.
Every day is a play day, and with so many
siblings and cousins around,
they keep themselves entertained, while the adults try to rest.
A cub can usually demand attention
and milk from any of the lactating mothers.
The mothers show irritation toward cubs that are not their own.
Going into the dry
winter,
the cubs grow,
along with their
demand for food.
The pressure increases for the lionesses to provide more meat.
And even the two Kakanaka brothers,
used to waiting on the sidelines, will soon be called
into
action.
Hunting requires a team effort,
and seasons of taking down buffalo have taught each lioness
exactly what she has to do.
But that expertise has made the Kakanakas dependent on a single food,
buffalo, despite
the other prey choices around them.
Giraffe hunting would demand a completely different strategy.
Giraffes, more alert than buffalo,
would spot the lions and their clumsy cubs from
far away.
The lions would have no chance to strike,
and giraffes aren't as gentle as they look.
They can deliver a kick that can knock a lion unconscious,
or even kill it.
So while buffalo are in the Kakanaka territory,
the lions concentrate on hunting them.
But the season has changed,
and as the buffalo exhaust each field of grass, they move to
Let's move to the next one.
Every day, a little further from the heart of the Cakanika Territory,
lions follow the herd,
constantly watching.
The females and cubs are here too, but hidden.
On their slow march,
the buffalo never notice more than one or two lions,
so they don't see the looming threat.
While the buffalo remain on the lion's turf, the pride shadows them,
keeping an eye on their next meal.
For several days,
the lions just follow the herd,
sizing up their options.
Choosing some suitable subjects, studying their movements,
and their positions within the herd.
Inevitably,
the time comes to attack.
One of the males and a lioness act as decoys.
They chase random victims to fragment the herd.
Unwittingly, the cubs get in the way, adding to the confusion.
But they're at risk of being trampled.
The second male
and the remaining females
attack the intended target.
Using all his strength, the male tries to suffocate it.
But the buffalo's straining muscles
form a steel barrier around its windpipe.
The lionesses do their part,
trying to immobilize it by breaking its spine.
Once he's down, he should be done.
But suddenly, the buffalo herd returns
to rescue their comrade,
confront the pride with their formidable armor.
They don't give up, lick the fallen animal
to encourage him to stand,
form a barrier against
the lions.
One of the buffalo challenges the predators,
and behind, their fallen fellow tries to rise.
As one charges again in front,
the buffalo that just moments ago was surrounded by killers
regains his footing and disappears into the herd.
For once, nature is kind to the prey,
rushes away, putting as much distance as possible
between them and their attackers.
Days of shadowing the herd have come to nothing.
Though weak with hunger and tired from the energetic attack,
the lions cannot rest.
They have to close in on the herd
and hunt again as soon as possible.
Lions despise water and won't cross unless their lives depend on it.
But they have no time to lose,
and the cubs have to keep pace with the adults.
Times are about to get harder.
The lions
are now approaching
the edge of their
20-square-mile territory.
It's the middle of winter.
And the buffalo follow the dwindling supply of greens
that fringe the rivers and channels of the Okavango.
It drives them further and further out,
to the limit of the Cakanika territory.
As the cold grip of winter draws
the lions' food source almost beyond their grasp,
the pride constantly looks for a chance to strike.
They take
a break.
Despite their hunger and the harsh weather,
the cubs seem almost untroubled.
A group roar warns surrounding prides that they hold this turf.
And today, they have something to shout about.
The five mothers
take time to bond
with their own cubs.
Reason for this pause
in their relentless pursuit lies hidden beneath a bush.
The sixth lioness has given birth
to two more cubs.
They swell the number of youngsters to an overwhelming 15.
Three mouths that need to be fed
while the Cakanika is at its most desperate.
And as the lion's main prey slips from their grasp,
buffalo leave, the elephants come.
Forced here when the edges of the Akavango go dry.
They seek Cakanika's water, though their food here is scarce.
They are less fussy eaters than the buffalo,
so they can make do with coarse grasses
and reeds the buffalo leave behind.
Trying to better their respective situations,
the two species cross paths.
One comes for the water.
The other leaves to seek sweet grasses.
And when the buffalo leave, the lions lose their main food source.
Elephants aren't on the menu. Not yet.
This is the border of the Cakanika territory,
an invisible line the lions won't cross.
Because another pride rules the other side.
But there's nothing to stop the buffalo from crossing.
Desperation drives the Cakanika pride to follow them over,
trespassing into enemy territory.
They don't get far before the locals spot them.
Cakanika female sizes up
the situation.
The lions forget their hunger for a while
as one of the Cakanika brothers prepares for a confrontation.
The buffalo can graze in peace while the cats settle their differences.
The Cakanikas make a stand.
Even though they are the intruders,
the other pride is at a disadvantage.
The dominant male is not with them,
so they turn back.
But the Kakonika male is not finished.
The resident pride makes a run for it.
Kakonika male
is too powerful.
The young males of the other pride can't stand up to him.
The larger Kakonika males assert
their dominance.
And though the resident pride retreats from the intruders,
they're still
the winners,
because their territory has
the buffalo herd.
The Kakonika intruders amble back to their own turf.
Despite all the posturing, the pride can't follow the buffalo.
Moving their cubs into enemy territory would be too dangerous.
As the lions return, the last of the buffalo leave
the Kakonika area for the rest of the season.
For the next six months, they'll share
the land with breeding herds of elephants.
The lion pride keeps moving, looking
for anything they can chase down
and eat.
They ignore the elephants,
because they're too large and difficult to hunt.
The elephants mostly ignore the lions as well.
As they try to glean up to 400 pounds of grass
and roots apiece from the depleted landscape,
the herd moves. Mothers guard
their youngest calves, small enough
to be attacked by predators.
The mothers dote on the babies for about four years.
Then they're on their own,
tasting independence for the first time.
Juvenile elephants risk wandering off and getting lost.
The hungry lions return to where the sixth lioness hid her cubs
when she joined the others to search for food.
Most times, mothers keep the hiding place
a secret from the rest of the pride.
The lioness brings the tiny additions,
now two months old, to meet their father.
But he's grumpy, so small
and weighing only nine pounds.
They will compromise the pride's movements.
Little steps won't be able to keep up.
Eventually, they will pay the price.
Already, some of the older cubs show signs of weakness.
They can't rest for long.
The lionesses take the lead.
The pride must travel far in search of food.
It takes a toll on the smallest and weakest members.
All they see
are large elephants.
They start to look
at them with interest.
With nothing else to eat, perhaps it's time to reconsider.
Too tired and weak to keep up with their
mothers, the cubs remain together, waiting.
They're on the brink of starvation.
More than half
of them have already died.
Fittest will have a fighting chance.
But without food, they'll have no chance at all.
Their mother's return brings no joy, just some affection.
The cubs try to drink, but the starving lioness's milk has gone dry.
Without milk or meat, the cubs' days are numbered.
When they move from mother to mother to try to suckle,
their desperate tugs irritate the lionesses.
At this point, even
the strongest
cubs only have
a few days to live,
hungry
and thirsty.
The cubs lie defeated.
The lionesses never give up.
They scan the horizon for something edible.
Nothing
but elephants.
Too large to hunt and traveling in tight groups.
But hunger calls.
The pride must hunt new prey.
Lionesses take their
chances, following the herd,
hoping for an opportunity.
Perhaps a youngster will get injured or stray from the herd.
The young elephants are unaccustomed to danger.
Eventually,
the lion's persistence pays off.
During this desperate time, nature gives the cats a lucky break.
But at the same time, it deals its cruelest hand.
A five-year-old youngster loses his way.
His trumpeting cries go unheard by his family.
He can't outrun the lions,
and his tusks are too small to use as weapons.
Lionesses drive the two-ton animal to the ground and hold him down.
Although the lionesses
are expert buffalo killers, they
have never tried to kill an elephant.
Their efforts are clumsy and ineffective.
Their teeth can't penetrate his one and a half inch thick skin,
and his neck is too wide to strangle.
So they attack the soft, vulnerable spots.
Without a muzzle to suffocate, they try to hold his trunk.
Unable to kill the elephant, they start to eat it alive.
First taste of meat after almost a month
of starvation revives their stamina,
and they fight for the right to every mouthful.
Twice the size of a buffalo,
the young elephant keeps the pride well-fed
and introduces them to a new kind of prey.
Over the next weeks, the lions work
to refine their elephant-killing techniques.
But the elephants strike back, stampeding
toward the lions at a ferocious pace.
They run for their lives.
The lions persist.
In this dry season that started so disastrously,
Killing elephants turns from an act of desperation
Do a way of life. Out of 15 cubs, seven survive.
They dine heartily with the eight adults.
Their success doesn't go unnoticed.
The neighboring pride, desperate for a meal, dares to confront them.
But the Okavango,
with a meal and their turf to protect,
won't tolerate it.
Chasing the intruders isn't enough.
They want to teach these trespassers a lesson.
Captured by the big male,
the young trespasser doesn't look him in the eyes.
A direct stare would mean
a challenge.
Too young to stand his ground against the powerful older male,
the intruding lion submits.
After failing to outrun him,
the damage is done.
The young male escapes with his life,
but he's branded forever, his genitals mutilated
in an ultimate act of dominance.
The other Okavango male asserts his own power
by having his way with the trespassing female.
By the end of winter,
under the washed-out skies
of an excruciatingly
hot October,
wave after wave of tired elephants move through the lion's territory.
Winter has taken its toll on the herds.
They've had their fill of life-giving water,
but the scrappy vegetation offers little
nutritional value. Mothers and calves grow weak.
And older calves stray from the herd and get lost.
Lionesses exploit
these opportunities to kill.
For the lions,
this has become a season of indulgence.
In one month, they kill five elephants,
ensuring their survival and teaching the cubs the ways of the hunt.
Now the cycle begins again.
It starts as a far-off flicker.
The darkness brings relief.
As soon as the drops hit the ground,
the Okavango comes alive and the animals return.
The breeding herds of elephants migrate back
to their summer feeding grounds,
where the rains will replenish the water
and revitalize their grazing areas.
Magically, the ruthlessness
of the past season gets washed away in a spray of green.
The most desperate season in decades.
About half the cubs survived.
A normal survival rate, even for this abnormally large pride.
Spread of fresh,
sweet grasses lures
the buffalo herds back into
the Okavango territory.
The lions eagerly watch their favorite prey arrive.
A ton of meat with a relatively soft hide to pierce,
and a short snout easy to smother.
The familiar game of cat
and mouse resumes,
though buffalo are no mice.
The buffalo boss aggressively approaches the lions,
hoping to frighten them away.
But the lions are too familiar with this prey.
During the next six months of rains,
the one-year-old cubs learn to hunt by trial
and error.
It's not as easy as their parents make it seem,
though they'll become experts soon enough.
But surviving
the harshest
season might have
made the cubs
overconfident.
Before they're ready,
they charge the buffalo.
Their inexperience puts their lives in jeopardy.
Buffalo are long-distance runners.
They can easily chase down the smaller, pesky lions and kill them.
Pride retreats, leaving an unfortunate cub in a precarious predicament.
He takes safety on a fallen tree stump.
However, he misjudges the height, putting him in the line of fire.
Bravely, he hangs on for his life.
Lions hunt together, but if one
gets in trouble, he's on his own.
Eventually, the buffalo lose interest,
and the pride returns to retrieve the shaken cub.
Pride puts
its skills
to the test
with every
hunt.
But with buffalo, they know the drill.
As usual, the adult females take the lead.
Attack the buffalo from the back to avoid the dangerous horns.
Pierce the hide and cling to the rump,
putting their entire weight on its back.
They try to pull the buffalo to the ground.
The cubs copy their mothers.
During the summer months, the constant supply
of buffalo meat builds the youngsters into strong 18-month-olds.
In six months,
they'll be self-sufficient killers.
By the end of the rainy season, the young lions walk confidently
with the adults, a hunting pride 14 strong.
The young males' manes have started to grow.
They follow their fathers' every move,
learning when to strike, when to wait,
when to hide, and when to go out in the open.
As sub-adults, they're old enough to test their strength
in one of the pride's most formidable battles.
They stride up front
with the two
Kakanaka brothers
and attack side by side.
The heavy male straddles the buffalo's back
and the female tries to trip it to the ground.
When the buffalo retaliate,
the cubs stand their ground with the adults.
The entire pride relentlessly attacks to weaken their prey.
The buffalo's leg breaks.
There's nothing his comrades can do for him now.
Again, the lions
attack from behind
to avoid the horns.
One of the males starts
to smother him.
With a broken leg, the buffalo fights for his life.
He stands,
and his comrades charge the lions.
Buffalo cannot walk.
Desperately wounded,
he wields his horns to inflict pain on his attackers.
Lesson for the younger lions
to learn.
In vain, he tries again to stand.
Lionesses and cubs continue their rear assault.
Keeping clear of the horns,
objective, the buffalo on his back,
so the male can finish him off.
A changing
landscape
of prosperity
and hardship,
the Kakanaka
pride has found
stability.
With the two strong brothers in command,
and a healthy number of youngsters to ensure the pride's existence,
they have come full circle.
From desperation sprang ingenuity, switching prey to win the battle,
waged by the harshest winter they've ever endured.
Life on the Akavango will always be unpredictable,
and the cubs will one day have to face that by themselves.
Their fathers
will expel them,
and they'll have to form prides of their own.
The struggle
of the season
is over,
but the struggle for survival never ends.
The Kakonikas will always be a pride in battle.